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Animals eat the plants, and ultimately the radiocarbon is distributed throughout the biosphere.
The ratio of λ is a constant that depends on the particular isotope; for a given isotope it is equal to the reciprocal of the mean-life – i.e.
The different elements of the carbon exchange reservoir vary in how much carbon they store, and in how long it takes for the These organisms contain about 1.3% of the carbon in the reservoir; sea organisms have a mass of less than 1% of those on land and are not shown on the diagram.
Accumulated dead organic matter, of both plants and animals, exceeds the mass of the biosphere by a factor of nearly 3, and since this matter is no longer exchanging carbon with its environment, it has a ratio having remained the same over the preceding few thousand years.
Other corrections must be made to account for the proportion of throughout the biosphere (reservoir effects).